The evaluation findings showed that the teachers perceived the PD to be valuable and relevant to their teaching practice—particularly the science content and the community building features of the PD. There were diverging opinions among the teachers about the value of the TSI pedagogical features. Teachers reported that they would continue to use the Exploring Our Fluid Earth curriculum website after the PD for activities and content and that they would continue to use the online learning community aspect of the website to interact with each other. Teachers’ self reports suggested that the PD helped them improve on their pedagogical content knowledge for teaching science (effect size = 0.24, p = .21), self-efficacy in teaching science through the process of inquiry (effect size = 1.50, p < .01), and metacognition in teaching (effect size = 0.18, p = .38). Teachers felt the PD was helpful in improving their ability to teach the science content, but some teachers found parts of the pedagogy to be difficult to implement. The teachers’ gains on the aquatic science content knowledge module-level assessments were consistently significantly higher in the post-test than in the pre-test (effect sizes ranged from 0.66 to 1.01; all p < .01); indicating the PD was successful in delivering content that was accessible to the teachers. Teachers reported having success implementing the content aspects of the target activities in their classrooms, and they improved in the quality of their implementation of TSI pedagogy as the project progressed. Teachers also improved in their understanding of the nature of inquiry-based science teaching (effect size = 1.24, p < .01); indicating that teachers matured in their breadth and depth of knowledge about teaching science through the process of inquiry. Students gained in their understanding of the nature-of-science (NOS) while participating in the project (in typical classrooms, effect size = 0.13, p < .05). Students of teachers who reported adhering more closely to the PD gained significantly more than the students in classes taught by teachers with lower adherence ratings (effect size = 0.21, p < .01); supporting the tentative conclusion that the PD had an effect on students’ NOS understanding. Students also gained in their content understanding (in typical classrooms, the effect size = 0.45, p < .01; in classrooms with teachers adhering more to the PD, there was an added effect size of 0.25, p < .01). High-school students did not gain as strongly in content as middle-school students (effect size = -0.34, p < .01). Students of teachers with little prior science PD experience showed significant gains in content knowledge (effect size = 0.18, p < .01), suggesting that the project benefited students in contexts with teachers that had less science PD exposure.