Biology Directory

This listing contains all faculty and associates of the Department of Biology. They may be affiliated with other programs or departments of UH Mānoa, but they are all active in the research, programs, and activities that the Department of Biology typically engages in.

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Rosanna Alegado

Assistant Researcher, Department of Oceanography and UH Sea Grant Program

Website

Email
Room

C-MORE 111

Phone

(808) 956-0565

Content Type: 
Faculty
Post Date: 
01/25/2016 at 9:59am
Last Modified: 
04/26/2016 at 1:00pm

Allen Allison

Senior Zoologist, Bishop Museum

Population Biology, Systematics, Herpetology, Papuan Herpetofauna

Email
Room

Bishop Museum

Phone

(808) 848-4145

Fax

(808) 847-8252

Content Type: 
Faculty
Post Date: 
10/09/2013 at 3:01pm
Last Modified: 
02/10/2017 at 4:39pm

Whitlow Au

Researcher Emeritus, Marine Mammal Research Program, Hawai'i Institute of Marine Biology

Echolocation, Hearing and Acoustics of Marine Mammals

Email
Room

HIMB

Phone

(808) 247-7426

Content Type: 
Faculty
Post Date: 
10/08/2013 at 12:39pm
Last Modified: 
04/27/2016 at 10:53am

Elizabeth Baker

Peer Advisor

Email
Room

EDM 216

Phone

(808) 956-5722

Fax

(808) 956-4745

Content Type: 
Staff
Post Date: 
11/30/2017 at 7:48am
Last Modified: 
11/30/2017 at 7:48am

Paul C. Banko

Faculty

Don't forget to enter a short version in the custom summary field (above).

Email
Room

Kilauea Field Station

Phone

(808) 967-7396

Fax

(808) 967-8568

Content Type: 
Faculty
Post Date: 
10/09/2013 at 3:08pm
Last Modified: 
04/27/2016 at 10:04am

Anthony Barley

Post-Doctoral Research Fellow

Phylogenetics, landscapes genetics, herpetology, conservation.

Email
Room

EDM 318A

Phone

(808) 956-9523

Fax

(808) 956-4745

Content Type: 
Faculty
Post Date: 
11/13/2014 at 9:35pm
Last Modified: 
11/13/2014 at 9:36pm

Kevin Bennett

Associate Professor

Associate Chair

I have a broad interest in structural and functional microscopy and diagnostic imaging, wth a focus on molecular imaging . My lab develops new contrast agents and imaging strategies to sensitively detect physiological and cellular function with magnetic resonance imaging. Our current projects range from discovery science to translational medicine.

Email
Room

EDM 309

Phone

(808) 956-8241

Fax

(808) 956-4745

Content Type: 
Faculty
Post Date: 
11/04/2017 at 9:41am
Last Modified: 
08/28/2018 at 1:03pm

Charles Birkeland

Unit Leader Emeritus, Hawaii Cooperative Fisheries Research Unit

Coral reef ecology and management, marine community ecology

Email
Room

N/A

Phone

N/A

Content Type: 
Faculty
Post Date: 
09/22/2017 at 8:21am
Last Modified: 
09/22/2017 at 8:21am

Brian Bowen

Researcher, Hawai'i Institute of Marine Biology

Phylogeography and Conservation Genetics of Marine Vertebrates

Website

Email
Room

HIMB

Phone

(808) 236-7426

Content Type: 
Faculty
Post Date: 
10/08/2013 at 12:49pm
Last Modified: 
05/10/2016 at 10:14am
Marguerite Butler

Marguerite Butler

Associate Professor

Functional Morphology, Biomechanics, Phylogenetics, Macroevolution, Modeling, and Statistics

Website

Email
Room

EDM 316

Phone

(808) 956-8303

Fax

(808) 956-4745

Content Type: 
Faculty
Post Date: 
09/02/2013 at 10:53pm
Last Modified: 
03/01/2017 at 3:54pm

Sean Callahan

Professor and Associate Chair

Prokaryotic Biology; "Fixation" of Biologically Inert Dinitrogen Gas and Specialized Cells Along a Filament

Email
Room

Sny 201

Phone

(808) 956-8015

Fax

(808) 956-5339

Content Type: 
Faculty
Post Date: 
10/10/2013 at 9:16am
Last Modified: 
04/25/2016 at 12:43pm

Christy Burt

Academic Support - Undergraduate Academic Advisor

Email
Room

EDM 216

Phone

(808) 956-6174

Fax

(808) 956-4745

Content Type: 
Staff
Post Date: 
08/21/2017 at 11:09am
Last Modified: 
08/21/2017 at 11:09am

Jessica Chen

Peer Advisor

Email
Room

EDM 216

Phone

(808) 956-8303

Fax

(808) 956-4745

Content Type: 
Staff
Post Date: 
11/02/2016 at 1:55pm
Last Modified: 
11/02/2016 at 1:55pm

Rebecca Chong

Assistant Professor

Evolution, Genomics, and Symbiosis

Email
Room

EDM 409

Phone

8089562512

Fax

TBA

Content Type: 
Faculty
Post Date: 
10/13/2017 at 9:26am
Last Modified: 
09/10/2018 at 6:46am
Kathleen S. Cole

Kathleen S. Cole

Associate Professor

Evolution of Behavior, Morphology and Developmental Biology

Email
Room

EDM 416

Phone

(808) 956-8618

Fax

(808) 956-4745

Content Type: 
Faculty
Post Date: 
09/09/2013 at 9:22pm
Last Modified: 
04/25/2016 at 12:32pm

Sheila Conant

Professor Emerita, Biology

Life History, Ecology and Conservation Biology of Endemic Hawaiian Birds, Island Biology

Email
Room

N/A

Phone

N/A

Content Type: 
Faculty
Post Date: 
09/22/2017 at 8:22am
Last Modified: 
09/22/2017 at 8:22am

Robert Cowie

Researcher, Center for Conservation Research and Training

The Origins and Determinants of Biological Diversity in the Islands of the Pacific

Website

Email
Room

Gil 408

Phone

(808) 956-4909

Fax

(808) 956-2647

Content Type: 
Faculty
Post Date: 
10/08/2013 at 1:22pm
Last Modified: 
05/03/2016 at 1:25pm
Curtis Daehler

Curtis Daehler

Professor

  • Population biology of invasive plants
  • Predicting plant invaders - Weed Risk Assessments
  • Global and regional patterns of plant invasions
  • Plant-herbivore interactions
  • Effects of fire on native and alien plant recruitment in coastal grasslands
Email
Room

St. John 413A

Phone

(808) 956-3929

Fax

(808) 956-3923

Content Type: 
Faculty
Post Date: 
10/10/2013 at 9:23am
Last Modified: 
04/25/2016 at 12:42pm
Heinz Gert de Couet

Heinz Gert de Couet

Professor

Biology Chair

Molecular Cell Biology and Developmental Evolution

Email
Room

EDM 413

Phone

(808) 956-9686

Fax

(808) 956-4745

Content Type: 
Faculty
Post Date: 
10/18/2017 at 7:45am
Last Modified: 
10/18/2017 at 7:45am

Julia DeJong

Office Assistant

Email
Room

EDM 216

Phone

(808) 956-8303

Fax

(808) 956-4745

Content Type: 
Staff
Post Date: 
09/21/2013 at 1:41pm
Last Modified: 
03/31/2017 at 9:33am

Pia Dizon

Program Manager

Email
Room

EDM 216

Phone

(808) 956-7315

Fax

(808) 956-4745

Content Type: 
Staff
Post Date: 
05/28/2015 at 12:09pm
Last Modified: 
05/28/2015 at 12:10pm

Megan Donahue

Associate Researcher, Hawai'i Institute of Marine Biology

Population Ecology, Particularly of Interspecific Interactions

Website

Email
Room

HIMB

Phone

(808) 236-7417

Fax

(808) 236-7443

Content Type: 
Faculty
Post Date: 
10/08/2013 at 1:31pm
Last Modified: 
04/26/2016 at 1:17pm

David Duffy

Professor, Department of Botany

How Natural Environments Respond to Pertubations

Email
Room

St. John 410

Phone

(808) 956-8218

Fax

(808) 956-4710

Content Type: 
Faculty
Post Date: 
10/08/2013 at 2:42pm
Last Modified: 
04/26/2016 at 1:19pm
Anuschka Faucci

Anuschka Faucci

Academic Support - Lab Coordinator

Email
Room

EDM 213

Phone

(808) 956-4742

Fax

(808) 956-4745

Content Type: 
Staff
Post Date: 
10/08/2013 at 11:29am
Last Modified: 
10/01/2014 at 11:20am

Alan Friedlander

Scientist, National Geographic Society

Coral Reef Community Ecology, Fisheries Science, Marine Conservation Biology, and Traditional Marine Resource Use and Management

Email
Room

Biomed A-103

Phone

(808) 956-8350

Fax

(808) 956-4238

Content Type: 
Faculty
Post Date: 
10/08/2013 at 2:49pm
Last Modified: 
04/27/2016 at 10:57am

Ruth Gates

Researcher and Director, Hawai'i Institute of Marine Biology

The Biology of Corals and the Evolution of Animal Sensory Systems

Website

Email
Room

HIMB Pauley 107

Phone

(808) 236-7420

Fax

(808) 236-7443

Content Type: 
Faculty
Post Date: 
10/08/2013 at 2:57pm
Last Modified: 
04/27/2016 at 10:21am

Samantha Giridhar

Institutional and Academic Support

Email
Room

EDM 216

Phone

(808) 956-4743

Content Type: 
Staff
Post Date: 
10/15/2014 at 10:42am
Last Modified: 
03/30/2017 at 8:22am

Michael G. Hadfield

Professor Emeritus, Pacific Biosciences Research Center

Larval Settlement and Metamorphosis; Conservation and Evolutionary Biology of Hawaiian Tree Snails

Email
Room

Henke 336/Kewalo Lab 214

Phone

(808) 956-6176

Content Type: 
Faculty
Post Date: 
09/30/2013 at 10:53pm
Last Modified: 
04/27/2016 at 10:58am

Daniel K. Hartline

Researcher and Director, Bekesy Lab

Neurophysiology and Ultrastructure of Crustacean Nervous Systems; Copepod Neurocology; Bioinformatic and Computational Studies of nerve Impulse Conduction

Email
Room

PBRC 119

Phone

(808) 956-8003

Fax

(808) 956-6984

Content Type: 
Faculty
Post Date: 
10/08/2013 at 3:17pm
Last Modified: 
01/31/2017 at 8:52am

Meihui He

Peer Advisor

Email
Room

EDM 216

Phone

(808) 956-5722

Fax

(808) 956-4745

Content Type: 
Staff
Post Date: 
11/30/2017 at 7:47am
Last Modified: 
11/30/2017 at 7:47am

Mark Hixon

Sidney & Erica Hsiao Endowed Professor of Marine Biology,

Chair of Zoology Graduate Program

Marine ecology and conservation biology, especially regarding coral reefs

Website

Email
Room

EDM 417

Phone

(808) 956-6427

Fax

(808) 956-4745

Content Type: 
Faculty
Post Date: 
11/13/2017 at 10:10am
Last Modified: 
09/17/2018 at 12:12pm

Kim N. Holland

Researcher, Hawai'i Institute of Marine Biology

Physiology and Behavior of Aquatic Animals; Resource Conservation

Website

Email
Room

HIMB

Phone

(808) 236-7410

Fax

(808) 236-7443

Content Type: 
Faculty
Post Date: 
09/06/2017 at 8:14am
Last Modified: 
09/06/2017 at 8:14am

Brenden Holland

Associate Researcher, Center for Conservation Research and Training

The Mechanisms and Processes that Result in Diversification of Multicellular Life on Earth

Email
Room

Henke 337

Phone

(808) 956-6176

Fax

(808) 956-2647

Content Type: 
Faculty
Post Date: 
10/09/2013 at 1:47pm
Last Modified: 
04/25/2016 at 2:46pm

Cynthia L. Hunter

Associate Professor, Biology

Director, Marine Option Program

Coral Reef Ecology and Conservation

Website

Email
Room

Dean 108/108 A

Phone

808-956-4748

Content Type: 
Faculty
Post Date: 
09/30/2013 at 11:21pm
Last Modified: 
05/30/2018 at 2:34pm

Kenneth Y. Kaneshiro

Director, CCRT/Director, HEBP/Research

Speciation, Evolutionary Biology, Sexual Selection Theory, Hawaiian Drosophilids

Email
Room

Gil 406

Phone

(808) 956-6739

Fax

(808) 956-2647

Content Type: 
Faculty
Post Date: 
10/08/2013 at 3:54pm
Last Modified: 
04/27/2016 at 10:35am

Robert A. Kinzie

Professor Emeritus, Biology

Coral ecology and photobiology; Aquatic ecology

Email
Room

N/A

Phone

N/A

Content Type: 
Faculty
Post Date: 
11/22/2013 at 9:47am
Last Modified: 
04/25/2016 at 2:01pm

Stephanie Kraft-Terry

Assistant Specialist - Academic Advising and Assessment, Interim Director of Advising

Academic Advising and Program-Level Assessment 

Email
Room

EDM 216 B

Phone

(808) 956-0735

Fax

(808) 956-4745

Content Type: 
Faculty
Post Date: 
08/09/2017 at 7:18am
Last Modified: 
03/14/2018 at 8:02am
Jeffrey Kuwabara

Jeffrey Kuwabara

Manoa Marine Option Program Coordinator

Email
Room

Dean 105A

Phone

(808) 956-8433

Fax

(808) 956-2417

Content Type: 
Staff
Post Date: 
11/21/2013 at 3:29pm
Last Modified: 
12/10/2013 at 3:43pm

Marc O. Lammers

Assistant Researcher, Hawai'i Institute of Marine Biology

Marine Bioacoustics and Cetacean Behavior

Website

Email
Room

HIMB

Phone

(808) 375-0010

Fax

(808) 236-7443

Content Type: 
Faculty
Post Date: 
09/06/2017 at 8:16am
Last Modified: 
08/28/2018 at 12:47pm

Petra Lenz

Researcher, Bekesy Lab

Neuroecology of Zooplankton Sensory Systems

Website

Email
Room

PBRC 119

Phone

(808) 956-8003

Fax

(808) 956-6984

Content Type: 
Faculty
Post Date: 
10/09/2013 at 2:44pm
Last Modified: 
02/10/2017 at 4:38pm

George S. Losey

Professor Emeritus, Biology

Ethology of reef fishes especially regarding UV vision

Email
Room

N/A

Phone

N/A

Content Type: 
Faculty
Post Date: 
11/22/2013 at 10:04am
Last Modified: 
05/03/2016 at 1:24pm

Matthew C. I. Medeiros

Assistant Research Professor

Evolutionary Biology, Ecology of Infectious Pathogens

Email
Room

PBRC, Bekesy Laboratory Rm 251

Phone

808-956-8187

Content Type: 
Faculty
Post Date: 
10/08/2013 at 4:00pm
Last Modified: 
01/19/2018 at 10:43am
Peter Marko

Peter Marko

Associate Professor

Evolution, Biogeography and Conservation

Website

Email
Room

EDM 317

Phone

(808) 956-6146

Fax

(808) 956-4745

Content Type: 
Faculty
Post Date: 
10/18/2017 at 7:44am
Last Modified: 
10/18/2017 at 7:44am

Margaret McFall-Ngai

Director, Pacific Biosciences Research Center

Professor, Kewalo Marine Laboratory

Animal-Bacterial Symbioses; the molecular 'design' of tissues that interact with light, including eyes and photophores

Email
Room

PBRC 215, Kewalo Lab 218

Phone

(808) 956-8838

Fax

(808) 956-4768

Content Type: 
Faculty
Post Date: 
04/25/2016 at 3:22pm
Last Modified: 
09/30/2016 at 1:15pm
Amy Moran

Amy Moran

Associate Professor

Marine Ecology and Evolution

Website

Email
Room

EDM 311

Phone

(808) 956-6147

Fax

(808) 956-4745

Content Type: 
Faculty
Post Date: 
09/30/2013 at 11:54pm
Last Modified: 
04/25/2016 at 12:35pm

Clifford W. Morden

Associate Professor

Plant Molecular Systematics, Evolution, Population Genetics, Hybrid Plant Biology, Biogeography, Conservation Biology

Email
Room

St. John 602A

Phone

(808) 956-9636

Fax

(808) 956-3923

Content Type: 
Faculty
Post Date: 
10/10/2013 at 9:49am
Last Modified: 
04/25/2016 at 12:46pm

Paul Nachtigall

Researcher and Director Emeritus, Marine Mammal Research Program, Hawai'i Institute of Marine Biology

Cetacean Sensory and Perceptual Processes

Website

Email
Room

HIMB

Phone

(808) 247-5297

Fax

(808) 247-5831

Content Type: 
Faculty
Post Date: 
08/28/2017 at 12:18pm
Last Modified: 
08/28/2017 at 12:18pm

Jain Pak

Academic Support - Undergraduate Academic Advisor

Email
Room

EDM 216

Phone

(808) 956-8996

Fax

(808) 956-4745

Content Type: 
Staff
Post Date: 
11/03/2016 at 7:53am
Last Modified: 
11/09/2016 at 11:17am

Megan Porter

Assistant Professor

Molecular Evolution; Visual System Evolution.

Email
Room

EDM 411

Phone

(808) 956-3266

Fax

(808) 956-4745

Content Type: 
Faculty
Post Date: 
01/15/2015 at 2:44pm
Last Modified: 
02/09/2018 at 7:08am

John Randall

Senior Ichthyologist Emeritus, Bishop Museum

Taxonomy and Biogeography of Tropical Marine Fishes

Email
Room

Bishop Museum

Phone

(808) 848-4130

Fax

(808) 847-8252

Content Type: 
Faculty
Post Date: 
10/09/2013 at 3:25pm
Last Modified: 
04/27/2016 at 11:18am
Floyd A. Reed

Floyd A. Reed

Associate Professor

An overall theme of my work is population genetics. I have interests in aspects of human evolution and in engineering safe and reversible gene drive systems.

Email
Room

EDM 412

Phone

(808) 956-6489

Fax

(808) 956-4745

Content Type: 
Faculty
Post Date: 
10/03/2017 at 8:18am
Last Modified: 
03/16/2018 at 8:52pm

Ernst S. Reese

Professor Emeritus, Biology

Behavioral ecology of coral reef animals

Email
Room

N/A

Phone

N/A

Content Type: 
Faculty
Post Date: 
11/22/2013 at 10:31am
Last Modified: 
05/03/2016 at 1:24pm

Robert H. Richmond

Research and Director, Kewalo Marine Laboratory

Marine Conservation, with a Focus on Coral Reefs

Email
Room

Kewalo Lab 218

Phone

(808) 539-7331

Fax

(808) 599-4817

Content Type: 
Faculty
Post Date: 
10/08/2013 at 4:12pm
Last Modified: 
04/26/2016 at 1:07pm

Ku'ulei Rodgers

Faculty

Mauka/Makai Connection Between Watersheds and Reefs Including Indicators of Reef Condition and Impacts of Ocean Acidification

Email
Room

HIMB

Phone

(808) 236-7440

Fax

(808) 236-7443

Content Type: 
Faculty
Post Date: 
10/09/2013 at 3:29pm
Last Modified: 
04/27/2016 at 10:05am
Andrew Rossiter

Andrew Rossiter

Director of State Aquarium

Director of Waikiki Aquarium

Awareness of the Ecology, Biology and Conservation Status of Fishes through Aquarium Exhibition

Email
Room

2777 Kalakaua Ave

Phone

(808) 923-9741

Fax

(808) 923-1771

Content Type: 
Faculty
Post Date: 
10/08/2013 at 9:45am
Last Modified: 
11/18/2013 at 10:16am

Edward G. Ruby

Professor, Kewalo Marine Lab

Microbe-host signaling; Bacterial behavior (bioluminescence, chemotaxis and quorum sensing); Comparative genomics and population biology

Email
Room

Kewalo Lab 318

Phone

(808) 539-7309

Fax

(808) 599-4817

Content Type: 
Faculty
Post Date: 
11/02/2016 at 2:53pm
Last Modified: 
05/10/2017 at 8:09am

Alison Sherwood

Graduate Chair

Algal Biodiversity

Email
Room

St. John 607C

Phone

(808) 956-3930

Fax

(808) 956-4115

Content Type: 
Faculty
Post Date: 
10/10/2013 at 9:57am
Last Modified: 
04/25/2016 at 12:45pm
Audrey Shintani

Audrey Shintani

Administration Officer

Email
Room

EDM 216

Phone

(808) 956-4711

Fax

(808) 956-4745

Content Type: 
Staff
Post Date: 
09/04/2013 at 9:34pm
Last Modified: 
05/28/2015 at 12:06pm

Celia Smith

Professor

Physiological Ecology of Reef Algae and Corals with Related Issues in Marine Ecology

Email
Room

St. John 614A

Phone

(808) 956-6947

Fax

(808) 956-3923

Content Type: 
Faculty
Post Date: 
10/10/2013 at 10:05am
Last Modified: 
04/25/2016 at 12:43pm
Cheryl Squair

Cheryl Squair

Academic Support - Lab Coordinator

Email
Room

EDM 213

Phone

(808) 956-4742

Fax

(808) 956-4745

Content Type: 
Staff
Post Date: 
10/08/2013 at 11:45am
Last Modified: 
10/01/2014 at 11:20am

John S. Stimson

Professor Emeritus, Biology

Population and community ecology of corals and reef fishes

Email
Room

N/A

Phone

N/A

Content Type: 
Faculty
Post Date: 
11/22/2013 at 10:39am
Last Modified: 
05/03/2016 at 1:25pm
Keao Sunaoka

Keao Sunaoka

Academic Support - Lab Coordinator

Email
Room

EDM 213

Phone

(808) 956-4742

Fax

(808) 956-4745

Content Type: 
Staff
Post Date: 
10/08/2013 at 11:51am
Last Modified: 
10/01/2014 at 11:21am
Andrew D. Taylor

Andrew D. Taylor

Associate Professor

Population and Community Ecology of Interspecific Interactions; Applied Statistics

Website

Email
Room

EDM 409

Phone

(808) 956-4706

Fax

(808) 956-4745

Content Type: 
Faculty
Post Date: 
10/03/2017 at 7:18am
Last Modified: 
10/03/2017 at 7:18am

Diana Thompson

Academic Support - Undergraduate Academic Advisor

Email
Room

EDM 216

Phone

(808) 956-0824

Fax

(808) 956-4745

Content Type: 
Staff
Post Date: 
04/27/2015 at 12:49pm
Last Modified: 
11/09/2016 at 11:27am
Robert Thomson

Robert Thomson

Associate Professor

Systematic Biology, Evolution and Conservation

Website

Email
Room

EDM 313

Phone

(808) 956-6476

Fax

(808) 956-4745

Content Type: 
Faculty
Post Date: 
10/03/2017 at 8:19am
Last Modified: 
01/30/2018 at 8:14am

Robert J. Toonen

Researcher, Hawai'i Institute of Marine Biology

Dispersal and Recruitment of Invertebrate Larvae; Population Genetics, Evolution & Ecology of Marine Invertebrates

Website

Email
Room

HIMB

Phone

(808) 236-7425

Fax

(808) 236-7443

Content Type: 
Faculty
Post Date: 
09/06/2017 at 8:15am
Last Modified: 
09/06/2017 at 8:15am

Les Watling

Professor

Biology of Deep-sea Octocorals and Their Associates, Biology of Crustacea, and Conservation Studies

Email
Room

EDM 312

Phone

(808) 956-8621

Fax

(808) 956-4745

Content Type: 
Faculty
Post Date: 
10/08/2013 at 10:30am
Last Modified: 
01/31/2017 at 8:55am
Christopher Z. Womersley

Christopher Z. Womersley

Professor

Physiological Ecology/Adaptive Biochemistry; Dehydration and Freezing Stress; Biological Control

Email
Room

Snyder Hall 406

Phone

(808) 956-8622; (808) 956-2264

Fax

(808) 956-4745

Content Type: 
Faculty
Post Date: 
10/08/2013 at 10:42am
Last Modified: 
04/25/2016 at 12:37pm

Joanne Yew

Assistant Researcher - Pacific Biosciences Research Center

Chemical communication of insects; neurobiology of chemosensory perception; development of mass spectrometry methods for natural product chemistry

Email
Room

Bekesy Lab 116

Phone

808-956-5870

Fax

808-956-6984

Content Type: 
Faculty
Post Date: 
05/08/2017 at 8:26am
Last Modified: 
05/18/2017 at 2:04pm
Masato Yoshizawa

Masato Yoshizawa

Assistant Professor

I am interested in the evolutionary mechanisms that generate novel "adaptive" function in animals.

Email
Room

EDM 414

Phone

(808) 956-6682

Fax

(808) 956-4745

Content Type: 
Faculty
Post Date: 
08/29/2014 at 11:01am
Last Modified: 
03/27/2018 at 10:51am